This installment of Sakura Taisen series takes place in the New York of 1928. For the first time in the series, you don't play as the brave young Ichirou Oogami, but as his nephew, Shinjiro Taiga. The young man is sent from Japan to serve in New York under the command of Ratchet Altair, the leader of the local Kagekidan (New York Combat Revue, Star Division), a group of young ladies who can pilot giant mechs and fight supernatural evil. A resurrection of a medieval Japanese warlord is about to take place in New York, and it's up to our heroes to stop the evil.